The grid is 0.1" holes because that's the spacing of most components; it's a bit like perfboard. If you're asking why they used a grid instead of a normal-style PCB, I'm not sure. It probably makes manufacturing the boards much easier since you can drill the holes with an assembly line rather than one at a time. The traces go between the holes; the traces are very narrow, so two traces can fit between a pair of holes. That's probably the tradeoff, that your traces need to be very precise and you probably need more layers because of all the holes in the way. The layout was probably done with CAD; PCBs with CAD go way back. IBM was doing circuit boards with CAD back in the early 1960s, using a flying spot of light to draw out the PCBs on photoresist.