Or simply put, can wild animals eat tomatoes safely(on evolutionary timescale) in winter if they don't normally grow in winter.
The tomato is native to places like Peru and Ecuador, and eventually was moved to [what is now known as] Mexico as we kept bringing it further north.
The tomato is Ohio's state fruit[1], but it does not belong in Ohio. The only reason we have tomatoes growing in Ohio is humans; nature has nothing to do with it. They wouldn't be here without people dilly-whacking with things.
(And I'm glad they did so. Tomatoes are delicious.)
[1]: Seems weird but it be that way anyhow. Ohio also has a native state fruit, which is the paw paw.