⌘R Reload in Safari; reply in Mail; reveal an alias's original in Finder
⌘L Address bar in browsers; message filter in Mail; clear to previous mark in Terminal.
⌘I File info in Finder; italics in text; inspector in Terminal.
I can keep going. This is not to say windows/osx is worse or better, just that claims of consistency are not suitable on either
What kind of consistency did you expect across Mail App and Safari? How do you "Reply" in browsers?