You can use dehumidifiers at a fraction of the power cost of full air heating, given that you only need to keep the surface pores of the e.g. wallpaper below a humidity that supports mold growth.
Similarly, the bathroom could gain a second insulated door-ish curtain for an airlock effect fairly easily.
I got a CO2 monitor, and noticed that it often went over 1000 ppm CO2. So I started cracking the windows slightly, which resolved it. There are downsides to sealing the house too tight.