Not knowledgable, but irradiated flies should not be expected to be irradiated again. There are 3 population pools:

1. The Factory spawning population - This is self-sustained, and never encounters radiation.

2. A subset of the spawned males from the factory population are irradiated, making them sterile.

3. The wild population, consisting of the sterile males + wild males + wild females.

If for some reason the sterile population is not fully sterile (unlikely), then maybe there is a gene that helps for radiation resistance, but the children of that strain will not encounter radiation, so it fades away.

The factories are not going out to the regions where the flies are deployed to get new fly studs.

I'd expect that if females start to breed more than once it would represent a problem. It's surprising it hasn't happened yet (for the ignorant of the field that I am at least)

I'm not sure radiation resistance is really a thing. Radiation causes physical damage, it's not like a virus or bacteria that an organism can potentially fight off with an immune response.

The few males that might survive the gamma exposure with intact fertility are probably just ones that didn't get a full dose.

It is rather amazing to me in fact that it's possible to sterilize the males without killing them.